The American Society for Cell Biology (ASCB) is an inclusive, international community of biologists studying the cell, the fundamental unit of life, and is dedicated to advancing scientific discovery, advocating sound research policies, improving education, promoting professional development, and increasing diversity in the scientific workforce. The first ASCB Annual Meeting was held November 2-4, 1961, in Chicago IL where 844 attendees gathered for three days of lectures, slides, and movies of cellular structure. Since then, the ASCB membership has grown to 9,000 worldwide, and now draw upwards of 5,000 people. Since 1960, 32 past or current ASCB members have won Nobel Prizes in medicine or in chemistry. This year, from December 3–6, 2017, the ASCB and the European Molecular Biology Organization (EMBO) are joining forces for the 2017 ASCB | EMBO Meeting, at the Pennsylvania Convention Center, 1101 Arch St, Philadelphia, PA 19107.